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In the honeybee, the three recombination hotspots are localized at the edges of three large inversions and they show a striking correlation with the large inversions in the honeybee genome [31]. In humans, recombination hotspots are correlated with large inversions, but they are usually much smaller than the ones observed in honeybees [47]. In humans, recombination hotspots are usually located close to inversion breakpoints, or in the vicinity of large deletions and duplications [48,49,50]. This is due to the fact that variation in recombination rate is not correlated with variation in the length of either inversions or deletions [41]. In humans, variation in recombination rate is caused by the presence of complex haplotypes or by the presence of genetic variants that alter recombination rates in cis [40,51,52,53]. Similar mechanisms are likely to be involved in honeybees, but they are not known yet. The three recombination hotspots in honeybees show a strong correlation with large inversions and large deletions (Fig. 2). This is even more remarkable in the case of the recombination hotspot at 39.

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